Chargaff's rules state that ().
A、in the DNA molecule the number of adenines = number of thymines and the number of guanines = number of cytosines
B、in the DNA molecule the number of guanines and cytosines = the number of adenines and thymines
C、the ratio of (A + T)/(G + C) is constant for a species
D、all the above
E、a and c only